Leaves: deltoid-ovate, oblong-ovate or lanceolate, 0.6–2 mm, margins recurved to erect distally, usually undifferentiated, apex obtuse or rounded, serrulate, with abaxial papillae, piliferous; costa excurrent, with an inconspicuous abaxial epidermis, filaments of 4–10 cells, cells cylindric to subspheric, [thick-walled], terminal cell rectangular, subspheric or conic with 2–5 solid papillae; cells of leaf base 11–70 µm, medial and distal cells 9–33 µm, smooth, thick-walled.

Sexual: condition gonioautoicous.

Seta: 5–20 mm.

Capsule: urn short-cylindric to ovoid-cylindric, 1–2.7 mm; operculum 0.6–1.1 mm; peristome short, nearly straight, cribose at base to long and strongly twisted, 198–1160 µm.

Spores: spheric, 9–22 µm.

Distribution: North America, nw Mexico, Europe, Asia (India), n Africa, Atlantic Islands (Canary Islands).

Discussion: Varieties 2 (2 in the flora).